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A Research
Report
By Dave Keehn
2005
was a year of firsts for the Georgia chapter. We pollinated our first
native Georgia trees with backcross pollen and harvested our first
nuts. The harvest yield was less than spectacular, but the experience
gained was invaluable. With that experience, and with some new trees
that we hope to pollinate, we look forward to a more fruitful year in
2006.
The nuts that we harvested this year are known as
‘BC3’, which means they represent the third backcross of a
Chinese-American tree with a pure American chestnut. These nuts are
roughly 95% American chestnut genetically, and will grow into mature
trees that are outwardly indistinguishable from a 100% American
chestnut tree. Genetically, their Chinese heritage provides some
level of chestnut blight resistance at this stage. Since the genetics
of blight resistance is not limited to a single gene, our work is not
done here.
In order to obtain
high levels of blight resistance, there are two more steps, called
intercrosses, in which we take BC3 trees and cross them with other
BC3 trees. These steps produce BC3F2 and finally BC3F3 trees. BC3F3
trees are the “finished product”, both characteristically American
chestnut in appearance and form, with blight resistance levels
comparable to a Chinese chestnut.
TACF Meadowview Research Farm produced the first BC3F3
nuts this year. However, in addition to these nuts being in very
limited supply, one of the tasks that each chapter is assigned is to
develop local “lines” of American chestnut from mother trees found in
the geographical area, in order to ensure maximum suitability for
eventual reforestation.
With the new Berry College orchard, we prepare for the
next step, which is to plant our BC3 trees in the ground. At Berry
College, we will be planting both the BC3 trees and some pure
American trees as controls.
The other challenge
that we face in restoring the American chestnut comes from the deadly
fungus Phytophora sp., known as black ink disease or root rot.
Phytophthora is particularly problematic in the Southeast. Long
before chestnut blight felled the American chestnut, phytophthora
devastated chestnut trees in the lower elevations of central,
southern and even coastal Georgia. Based on field observations, we
have recognized that the southern chapters also need to breed for
phytophthora resistance. |